Peter Weston had previously published Speculation and Nexus in the 1960s and 1970s. For his return to fanzine publishing, he originally launched Prolapse in 1983 as an apazine. In 2006, it was relaunched, and featured letters of comment on the previous two issues. With the fourth issue, Weston announced its focus from that point forward would be the history of British fandom. It was subtitled "The Time-Travelling Fanzine".
Highlights include the many older fans who contribute, such as Brian Aldiss, Malcolm Edwards (Vector), Harry Harrison, Jim Linwood (Typo), George Locke (Smoke), Mike Moorcock (A Fanzine Called Eustace), and Christopher Priest.
Other contributors include Judy Blish, Bruce Burn, Chuck Connor (IDOMO), Dick Ellingsworth, Rob Hansen (Chuch), Ann Green (Ormolu), John Hall, John-Henri Holmberg, Dave Jenrette, Tony Keen, Dan Morgan, Peter Nicholls, Stan Nicholls (Gothique), Greg Pickersgill (Stop Breaking Down, Rastus Johnson's Cakewalk), Mark Plummer (Banana Wings), and Mark Tudor.
Art work was contributed by Dave Hicks, Sam Long, and Andrew Stephenson.
Photography was contributed by Dave Barber, Chris Bell, Simon Bradshaw, Bill Burn, Bill Burns, Ramsey Campbell, Les Flood, Keith Freeman, Rob Hansen (Chuch), John Harvey, Dick Howett, Terry Jeeves (ERG, The Damned Patrol), Peter Mabey, Mike Meara, Beryl Mercer (Link, Oz), Sam Mercer, Ellis Mill, Chris Murphy, Harry Nadler, Robert Quaglia, Ina Shorrock, Norman Shorrock, Ian Sorenson, and Lars-Olav Strangberg and Peter Weston.
Also included was an extensive letter column with long letters from Brian Aldiss, John Berry (Pot Pourri, The Damned Patrol), Doug Bell (Head), Harry Bell (Grimwab), Sandra Bond (QuasiQuote), Claire Brialey (Journey Planet), Randy Byers (Chunga), Graham Charnock, Chuck Connor (IDOMO), Lilian Edwards (Floss!, Gloss, This Never Happens), Malcolm Edwards (Vector), Steve Green, Joseph Nicholas (Vector), Darroll Pardoe (Les Spinge), Rosemary Pardoe (Wark, Ghosts & Scholars), Joe Patrizio (Ipso Facto), Charles Platt (Beyond), Mark Plummer (Banana Wings), Doreen Rogers, Steve Stiles, Simone (Walsh) Restall (Seamonsters), Lloyd Penney (Torus), and Peter Roberts (Checkpoint).